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Overall, if you average these performances the PCIe x4 slot got % the performance of the PCIe x16 slot There may be other reasons

4x4 super slot slot and a slot they call a pcie video slot its 4x it also has pci slots for my older uad-1s really a weirdo mobo i asked and asked before getting a uad

4x4 joker slot In ASUS 「PCIe x16 Slots - 4」 - Motherboards Components , you could find most 4 x 5 x SlimSAS SATA 6Gbs 1-4